This species is included in the Betta unimaculata complex of closely-related species within the genus, of which members share the following set of characters: body long and slender with depth at dorsal fin origin 18-25 % SL; head large and blunt with width 19-24 % SL; long maxilla and lower lip with distance from tip of lower jaw to posterior end of maxilla 27-54 % HL; caudal-fin rounded in shape, occasionally with elongated median rays; pelvic-fin short and filamentous; dorsal and anal fins relatively pointed.

This species was previously included in the Betta foerschi group of closely-related species within the genus, but following its redescription in 2012 now lends its name to the B. rubra group.

It was first placed into its own species group by Witte and Schmidt (1992), primarily on the basis of a triangular marking that appears below the eyes of the fish, although Tan and Ng (2005) suggested that this feature is insufficient to warrant the fo…

B. channoides and B. albimarginata together comprise the Betta albimarginata group/complex of closely-related species within the genus, these differing from all other members of the genus in colour pattern and by possession of 9-12 anal-fin spines (vs. 0-4). They look very similar to one another but differ in the following characters: anal fin spines 9-11 in B. albimarginata vs. 12 in B. channoides; anal fin rays 21-23 vs. 23-25; lateral scale rows 26-27 vs. 27.5-28; trans…

This species is often confused with B. krataios in the aquarium trade, the latter having been known as B. sp. ‘Kapuas’ prior to description. The two can easily be told apart as the caudal-fin is rounded in B. krataios but spade-shaped and in adults, highly-extended in B. dimidiata.

Together these two comprise the B. dimidiata group of closely-related species within the gen…

This species is included in the B. bellica group/complex of closely-related species within the genus, an assemblage of which members share the following set of characters: long and slender body with dorsal and ventral margins almost parallel; body depth 23-28 % SL; 30-33 anal-fin rays; 11-13 dorsal-fin rays; 32-34 total vertebrae; body dark brown in colour with iridescent green markings on each individual scale.

At one locality close to Serian, Sarawak, the habitat comprised a swiftly-flowing, clear hill stream with a substrate of quartz gravel and variably-sized rocks. The pH was 7.0 and there was no aquatic vegetation. The stream was approximately 10 metres across at its widest point and between 5-80 cm deep. B. taeniata was found in sluggish stretches among submerged tree roots, leaf litter, or overhanging riparian vegetation.

B. falx is included in the Betta picta group/complex of closely-related species within the genus, an assemblage comprising species with the following shared characters: unpaired fins rounded (anal-fin sometimes tapering to a blunt point); anal-fin rays I-III, 18-24; dark marginal band on anal and caudal fins, more pronounced in mature males; presence of pre-orbital and post-orbital stripes; presence of chin bar on throat; iridescent green, gold, or blue scales on opercle.

It should be noted that not all authors consider the B. pugnax group to contain the same species, with some placed in the B. picta complex by Schindler and Schmidt (2006), for example.

B. raja can be told apart from other members of the B. pugnax group by the following characters: mature males with very long pelvic fins covering 15-23 anal-fin rays or even longer than the base of the anal-fin in some cases; anal-fin and lower caudal-fin with black marginal bands; 25-28 anal-fin rays; 30-32 lat…

B. macrostoma is included in the Betta unimaculata complex of closely-related species within the genus, of which members share the following set of characters: body long and slender with depth at dorsal fin origin 18-25 % SL; head large and blunt with width 19-24 % SL; long maxilla and lower lip with distance from tip of lower jaw to posterior end of maxilla 27-54 % HL; caudal-fin rounded in shape, occasionally with elongated median rays; pelvic-fin short and filamentous; dorsal and anal fins relatively pointed.

This species was traded as B. sp. ‘Kapuas’ prior to its formal description and continues to be mislabelled B. dimidiata. The two can easily be told apart as the caudal-fin is rounded in B. krataios but spade-shaped and in adults, highly-extended in B. dimidiata.

Together these two comprise the B. dimidiata group/complex of closely-related species within the genus, members of which can be characterised b…