N. binotatus is distinguished from similar-looking congeners such as N. longistriatus and N. ornatus by its unique body patterning consisting very simply of two dark stripes, one running along the dorsal surface and the other from the tip of the snout, through the eye and along the flank.

There are usually 1-3 small, vertically-orientated spots extending above the lateral stripe between the pectoral-fin and dorsal fins. These are su…

Fishes which inhabit similar biotopes in nature constitute the best options, especially peaceful, open water-dwelling cyprinids since the presence of one or two schools can make a visible difference to the confidence of this naturally reclusive loach.

Other possibilities include rheoph…

Among them it’s most similar to I. pumila which was initially described as a subspecies but is now considered distinct. The two are most easily-separated by colour pattern; I. koreensis has 10 or…

It’s most easily told apart from K. brevifasciata, its only congener, by body patterning consisting of dark, vertical bars running along the dorso-ventral part of the body (vs. dark bars running alongside the midventral part of the body), lips with wrinkled surfaces (vs. smooth) and barbels short (vs. long).

Patterning is highly variable depending both within and between populations and the species may comprise an assemblage containing a number of taxa (Kottelat and Lim, 1993; Tan and Kottelat, 2009). For example, those from Singapore and Johor possessing intricate mottling on the dorsal surface while individuals from Terengganu have a series of saddle-like markings either alternating or connecting with the midlateral markings.

It’s included in the P. kuhlii–oblonga group of closely-related species within the genus as per Bohlen et al. (2011) and can be…

Some Sumatran and Bornean populations of the highly variable P. shelfordii superficially resemble P. piperata which can cause issues with identification as the two often occur together. In general however the body patterning in P. piperata is than that of P. shelfordii and they differ in vertebral counts (mode 47 vs. 50, respectively).

It’s included in the P. kuhlii group of closely-related species and can be distinguished from other members by the following combination of characters: 8-11 very broad, quadrangular, regular dark body bars; orange base body colour; caudal-fin either entirely black or with a large black blotch at the base and a submarginal dark bar or row of spots. The latter feature can be useful in identifying young or newly-imported, undernourished specimens which can…

Originally known only from Johor and Pahang states, Peninsular Malaysia but since recorded from the Batang Hari river drainage, Sumatra and the Kapuas system, Kalimantan Barat (West Kalimantan) province, Indonesian Borneo. Type locality is ‘Tahan River, Pahang, Malaysia’.

This species was available in the hobby several years prior to being described, invariably traded as G. sp. aff. altifrons ‘Tocantins’, G. sp. ‘Tocantins’ or G. sp. ‘rio Areoes’. The latter of these names refers to an Indian reserve named ‘Areões’ in the rio Das Mortes drainage, a tributary of the Araguaia which itself drains into the Tocantins, and the aquarium specimens in our images have been identified as G. neambi by Dr. Paulo Lucinda, co-describer of the species.